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Mark Greenwood

Mark Greenwood

PhD Student

Sainsbury Laboratory
University of Cambridge
47 Bateman Street

Research Interests

Broadly, I am interested in how cells work together to create useful systems. In my PhD in James Locke’s group, I am using a mixture of experiments and theory to understand how cells interact to keep track of the time of day. We found that the circadian clock in individual cells of Arabidopsis thaliana can use local cell-to-cell signalling to agree on the time. I now hope to further this work towards an understanding that may inform growth strategies in the field.

Prior to my PhD I studied for my undergraduate degree in Genetics and Masters in Post-Genomic Sciences at the University of Liverpool. During this time I worked with Anthony Hall who peaked my interest in plants. I also had the opportunity to complete other varied research projects, including in engineering at McGill University and Synthetic Biology at Synthace, London.

Key Publications

Greenwood M, Domijan M, Gould PD, Hall AJW, Locke JCW. Coordinated circadian timing through the integration of local inputs in Arabidopsis thaliana. PLoS Biol. 2019;17: e3000407. (forthcoming)


Rees H, Duncan S, Gould P, Wells R, Greenwood M, Brabbs T, et al. A high-throughput delayed fluorescence method reveals underlying differences in the control of circadian rhythms in Triticum aestivum and Brassica napus. Plant Methods. B2019;15: 51.


Gould PD, Domijan M, Greenwood M, Tokuda IT, Rees H, Kozma-Bognar L, et al. Coordination of robust single cell rhythms in the Arabidopsis circadian clock via spatial waves of gene expression. eLife. eLife Sciences Publications Limited; 2018;7: e31700. doi:10.7554/eLife.31700


Tindall AJ, Waller J, Greenwood M, Gould PD, Hartwell J, Hall A. A comparison of high-throughput techniques for assaying circadian rhythms in plants. Plant Methods. 2015;11: 32.


Greenwood M, Langlois R, Waters KE. The potential for dry processing using a Knelson Concentrator. Miner Eng. 2013;45: 44–46. 

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